In 1926, a time of great fear after a destructive war between humans and demons, the mutating monsters known as the Kouma threaten Tokyo. Led by Maria, Sumire, Leni and, of course, Sakura, the elite combat force known as the Flower Division is charged with defending Tokyo. However, the force is further threatened by the arrival of a nefarious U.S. company that is attempting to market a new defense weapon, and complicated by the arrival of Lachette, a member of the troupe when it was the Star division.

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